Ishihara, H. & Ishiyama, R. (1986)
Systematics and distribution of the skates of the north Pacific. In Uyeno, T. & Arai, R. & Taniuchi, T. & Matsuura, K. (Eds) Proceedings of the second international conference on Indo-Pacific fishes. Ichthyological Society of Japan, Tokyo: 269–280
McEachran, J.D. & Miyake, T. (1986)
Interrelationships within a putative monophyletic groups of skates (Chondrichthyes, Rajoidei, Rajini). In Uyeno, T. & Arai, R. & Taniuchi, T. & Matsuura, K. (Eds) Proceedings of the second international conference on Indo-Pacific fishes. Ichthyological Society of Japan, Tokyo: 281–290
Deynat, P.P. & Séret, B. (1996)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). I- Morphologie et arrangement des denticules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 17(2), 65–83
Camhi, M.D. & Fowler, S. & Musik, J. & Brautigam, A. & Fordham, S.V. (1998)
Sharks and their relatives. Ecology and conservation. Occasional papers of the IUCN species survival commission, 20, 39p.
Deynat, P.P. (1998)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). II. Morphologie et arrangement des tubercules cutanés . Annales des Sciences Naturelles – Zoologie et Biologie Animale, 19(3–4), 155–172
DOI: 10.1016/S0003-4339(99)80004-5