Gordon, J.D.M. & Duncan, J.A.R. (1989)
A Note on The Distribution and Diet of Deep-Water Rays (Rajidae) in An Area of The Rockall Trough. Journal of the Marine Biological Association of the United Kingdom, 69(3), 655–658
DOI: 10.1017/S0025315400031040
Deynat, P.P. & Brito, P.M.M. (1994)
Révision des tubercules cutanés de raies (Chondrichthyes, Batoidea) du Bassin du Paraná, tertiaire d'Amérique du Sud. Annales de Paléontologie, 80(4), 237–251
McEachran, J.D. & Konstantinou, H. (1996)
Survey of the variation of alar and malar thorns in skates: phylogenetic implications (Chondrichthyes: Rajoidei). Journal of Morphology, 228(2), 165–178
DOI: 10.1002/(SICI)1097-4687(199605)228:2<165::AID-JMOR5>3.0.CO;2-4