Stehmann, M. (1970)
Vergleichend morphologische und anatomische Untersuchungen zur Neuordnung der Systematik der nordostatlantischen Rajidae (Chondrichthyes, Batoidei). Archiv für Fischereiwissenschaft, 21(2), 73–164

Lloris, D. (1986)
Ictiofauna demersal y aspectos biogeográficos de la costa Sudoccidental de Africa (SWA/NAMIBIA). Monografías de zoología marina, Instituto de Ciencias del Mar, Barcelona, Consejo Superior de Investigaciones Científicas, 1: 429pp

McEachran, J.D. & Miyake, T. (1986)
Interrelationships within a putative monophyletic groups of skates (Chondrichthyes, Rajoidei, Rajini). In Uyeno, T. & Arai, R. & Taniuchi, T. & Matsuura, K. (Eds) Proceedings of the second international conference on Indo-Pacific fishes. Ichthyological Society of Japan, Tokyo: 281–290

Bauchot, M.-L. (1987)
Requins, Raies et autres Batoides, Chimeres in: Fischer W., Schneider M & M.–L. Bauchot(eds.): Fiches FAO d'identification des especes pour les besoins de la peche.(Revision l). Mediterranee et Mer Noire. Zone de peche 37. Vol. 2 Vertebres. Rome, CEE, FAO:: 767–890

Compagno, L.J.V. & Ebert, D.A. & Smale, M.J. (1989)
Guide to the sharks and rays of Southern Africa. New Holland Ltd., London 1989

Compagno, L.J.V. & Ebert, D.A. & Cowley, P.D. (1991)
Distribution of offshore demersal cartilaginous fishes (Class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11(1), 43–139, figs 1–24
DOI: 10.2989/025776191784287664

Deynat, P.P. & Séret, B. (1996)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). I- Morphologie et arrangement des denticules cutanés. Annales des Sciences Naturelles – Zoologie et Biologie Animale, 17(2), 65–83

Lanfranco, G.G. (1996)
The fish around Malta (Central Mediterranean). Progress Press Co. Ltd.(Malta). 132 p.

McEachran, J.D. & Konstantinou, H. (1996)
Survey of the variation of alar and malar thorns in skates: phylogenetic implications (Chondrichthyes: Rajoidei). Journal of Morphology, 228(2), 165–178
DOI: 10.1002/(SICI)1097-4687(199605)228:2<165::AID-JMOR5>3.0.CO;2-4

Deynat, P.P. (1998)
Le revêtement cutané des raies (Chondrichthyes, Elasmobranchii, Batoidea). II. Morphologie et arrangement des tubercules cutanés . Annales des Sciences Naturelles – Zoologie et Biologie Animale, 19(3–4), 155–172
DOI: 10.1016/S0003-4339(99)80004-5

Borges, T.C. & Costa, E. & Cristo, M. & Erzini, K. & Malaquias, A. & Nortista, P. & Olim, S. & Pais, C. & Sendão, J. & Campos, A. & Fonseca, P. & Santos, J. & Larsen, R. & Eide, A. & Broadhurst, M. (2002)
Managing By-Catch and Discards: A Multidisciplinary Approach ("Bydiscard"). Study Project No.99/058 Final Report to the European Commission Directorate General Fisheries 2002

Relini, G. & Mannini, A. & Piano, T. (2003)
Annotated checklist of the skates (Chondrichthyes, Rajidae) in the Northern Ligurian Sea. Biologia Marina Mediterranea, 10(2), 882–885

Farrag, M.M.S. & El-Naggar, H.A. & Abou-Mahmoud, M.M.A. & Alabssawy, A.N. & Ahmed, H.O. & Abo-Taleb, H.A. & Kostas, K. (2019)
Marine biodiversity patterns off Alexandria area, southeastern Mediterranean Sea, Egypt. Environmental Monitoring and Assessment, 191(6), 367
DOI: 10.1007/s10661-019-7471-7