Lebedev, O.A. (1995)
Middle Famennian (Upper Devonian) Chondrichthyans and Sarcopterygians from Oryol region; Central Russia. Geobios, 28(Supplement 2), 361–368
DOI: 10.1016/S0016-6995(95)80139-1

Lebedev, O.A. & Ivanov, A.O. & Linkevich, V.V. (2020)
Chondrichthyan spines from the Famennian (Upper Devonian) of Russia. Acta Geologica Polonica, 70(3), 339–362
DOI: 10.24425/agp.2020.132255